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V. Mutual Affinities of Organic Beings: MorphologyEmbryologyRudimentary Organs Analogical Resemblances WE can understand why a species or a group of species may depart from its allies, in several of its most important characteristics, and yet be safely classed with them. This may be safely done, and is often done, as long as a sufficient number of characters, let them be ever so unimportant, betrays adventure tour travel agency hidden bond of community of descent. Let two forms have not a single character in common, yet, if these extreme forms are connected together by a chain of intermediate groups, we may at once infer their community of descent, and we put them all into adventure tour travel agency same class. As we find organs of high physiological importancethose which serve to preserve life under adventure tour travel agency most diverse conditions of existenceare generally adventure tour travel agency most constant, we attach especial value to them; but if these same organs, in another group or section of a group, are found to differ much, we at once value them less in our classification. We shall presently see why embryological characters are of such high classificatory importance. Geographical distribution may sometimes be brought usefully into play in classing large genera, because all adventure tour travel agency species of adventure tour travel agency same genus, inhabiting any distinct and isolated region, are in all probability descended from adventure tour travel agency same parents. 1 We can understand, on adventure tour travel agency above views, adventure tour travel agency very important distinction between real affinities and analogical or adaptive resemblances. Lamarck first called attention to this subject, and he has been ably followed by Macleay and others. The resemblance in adventure tour travel agency shape of adventure tour travel agency body and in adventure tour travel agency fin-like anterior limbs between dugongs and whales, and between these two orders of mammals and fishes, are analogical. So is adventure tour travel agency resemblance between a mouse and a shrewmouse (Sorex), which belong to different orders; and adventure tour travel agency still closer resemblance, insisted on by Mr. Mivart, between adventure tour travel agency mouse and a small marsupial animal (Antechinus) of Australia. These latter resemblances may be accounted for, as it seems to me, by adaptation for similarly active movements through thickets and herbage, together with concealment from enemies. 2 Amongst insects there are innumerable similar instances; thus Linnæus, misled by external appearances, actually classed an homopterous insect as a moth. We see something of adventure tour travel agency same kind even with our domestic varieties, as in adventure tour travel agency strikingly similar shape of adventure tour travel agency body in adventure tour travel agency improved breeds of adventure tour travel agency Chinese and common pig, which are descended from distinct species; and in adventure tour travel agency similarly thickened stems of adventure tour travel agency common and specifically distinct Swedish turnip. The resemblance between adventure tour travel agency greyhound and adventure tour travel agency race-horse is hardly more fanciful than adventure tour travel agency analogies which have been drawn by some authors between widely different animals. 3 On adventure tour travel agency view of characters being of real importance for classification, only in so far as they reveal descent, we can clearly understand why analogical or adaptive characters, although of adventure tour travel agency utmost importance to adventure tour travel agency welfare of adventure tour travel agency being, are almost valueless to adventure tour travel agency systematist. For animals, belonging to two most distinct lines of descent, may have become adapted to similar conditions, and thus have assumed a close external resemblance; but such resemblances will not revealwill rather tend to conceal their blood-relationship. We can thus understand adventure tour travel agency apparent paradox, that adventure tour travel agency very same characters are analogical when one group is compared with another, but give true affinities when adventure tour travel agency members of adventure tour travel agency same group are compared together: thus, adventure tour travel agency shape of adventure tour travel agency body and fin-like limbs are only analogical when whales are compared with fishes, being adaptations in both classes for swimming through adventure tour travel agency water; but between adventure tour travel agency several members of adventure tour travel agency whale family, adventure tour travel agency shape of adventure tour travel agency body and adventure tour travel agency fin-like limbs offer characters exhibiting true affinity; for as these parts are so nearly similar throughout adventure tour travel agency whole family, we cannot doubt that they have been inherited from a common ancestor. So it is with fishes. 4 Numerous cases could be given of striking resemblances in quite distinct beings between single parts or organs, which have been adapted for adventure tour travel agency same functions. A good instance is afforded by adventure tour travel agency close resemblance of adventure tour travel agency jaws of adventure tour travel agency dog and Tasmanian wolf or Thylacinus,animals which are widely sundered in adventure tour travel agency natural system. But this resemblance is confined to general appearance, as in adventure tour travel agency prominence of adventure tour travel agency canines, and in adventure tour travel agency cutting shape of adventure tour travel agency molar teeth. For adventure tour travel agency teeth really differ much: thus adventure tour travel agency dog has on each side of adventure tour travel agency upper jaw four pre-molars and only two molars; whilst adventure tour travel agency Thylacinus has three pre-molars and four molars. The molars also differ much in adventure tour travel agency two animals in relative size and structure. The adult dentition is preceded by a widely different milk dentition. Any one may of course deny that adventure tour travel agency teeth in either case have been adapted for tearing flesh, through adventure tour travel agency natural selection of successive variations; but if this be admitted in adventure tour travel agency one case, it is unintelligible to me that it should be denied in adventure tour travel agency other. I am glad to find that so high an authority as Professor Flower has come to this same conclusion. 5 The extraordinary cases given in a former chapter, of widely different fishes possessing electric organs,of widely different insects possessing luminous organs,and of orchids and asclepiads having pollen-masses with viscid discs, come under this same head of analogical resemblances. But these cases are so wonderful that they were introduced as difficulties or objections to our theory. In all such cases some fundamental difference in adventure tour travel agency growth or development of adventure tour travel agency parts, and generally in their matured structure, can be detected. The end gained is adventure tour travel agency same, but adventure tour travel agency means, though appearing superficially to be adventure tour travel agency same, are essentially different. The principle formerly alluded to under adventure tour travel agency term of analogical variation has probably in these cases often come into play; that is, adventure tour travel agency members of adventure tour travel agency same class, although only distantly allied, have inherited so much in common in their constitution, that they are apt to vary under similar exciting causes in a similar manner; and this would obviously aid in adventure tour travel agency acquirement through natural selection of parts or organs, strikingly like each other, independently of their direct inheritance from a common progenitor. 6 As species belonging to distinct classes have often been adapted by successive slight modifications to live under nearly similar circumstances,to inhabit, for instance, adventure tour travel agency three elements of land, air, and water,we can perhaps understand how it is that a numerical parallelism has sometimes been observed between adventure tour travel agency sub-groups of distinct classes. A naturalist, struck with a parallelism of this nature, by arbitrarily raising or sinking adventure tour travel agency value of adventure tour travel agency groups in several classes (and all our experience shows that their valuation is as yet arbitrary), could easily extend adventure tour travel agency parallelism over a wide range; and thus adventure tour travel agency septenary, quinary, quarternary and ternary classifications have probably arisen. 7 There is another and curious class of cases in which close external resemblance does not depend on adaptation to similar habits of life, but has been gained for adventure tour travel agency sake of protection. I allude to adventure tour travel agency wonderful manner in which certain butterflies imitate, as first described by Mr. Bates, other and quite distinct species. This excellent observer has shown that in some districts of S. America, where, for instance, an Ithomia abounds in gaudy swarms, another butterfly, namely, a leptalis, is often found mingled in adventure tour travel agency same flock; and adventure tour travel agency latter so closely resembles adventure tour travel agency Ithomia in every shade and stripe of colour and even in adventure tour travel agency shape of its wings, that Mr. Bates, with his eyes sharpened by collecting during eleven years, was, though always on his guard, continually deceived. When adventure tour travel agency mockers and adventure tour travel agency mocked are caught and compared, they are found to be very different in essential structure, and to belong not only to distinct genera, but often to distinct families. Had this mimicry occurred in only one or two instances, it might have been passed over as a strange coincidence. But, if we proceed from a district where one Leptalis imitates an Ithomia, another mocking and mocked species, belonging to adventure tour travel agency same two genera, equally close in their resemblance, may be found. Altogether no less than ten genera are enumerated, which include species that imitate other butterflies. The mockers and mocked always inhabit adventure tour travel agency same region; we never find an imitator living remote from adventure tour travel agency form which it imitates. The mockers are almost invariably rare insects; adventure tour travel agency mocked in almost every case abound in swarms. In adventure tour travel agency same district in which a species of laptalis closely imitates an Ithomia, there are sometimes other Lepidoptera mimicking adventure tour travel agency same Ithomia: so that in adventure tour travel agency same place, species of three genera of butterflies and even a moth are found all closely resembling a butterfly belonging to a fourth genus. It deserves especial notice that many of adventure tour travel agency mimicking forms of adventure tour travel agency leptalis, as well as of adventure tour travel agency mimicked forms, can be shown by a graduated series to be merely varieties of adventure tour travel agency same species; whilst others are undoubtedly distinct species. But why, it may be asked, are certain forms treated as adventure tour travel agency mimicked and others as adventure tour travel agency mimickers? Mr. Bates satisfactorily answers this question, by showing that adventure tour travel agency form which is imitated keeps adventure tour travel agency usual dress of adventure tour travel agency group to which it belongs, whilst adventure tour travel agency counterfeiters have changed their dress and do not resemble their nearest allies. 8 We are next led to inquire what reason can be assigned for certain butterflies and moths so often assuming adventure tour travel agency dress of another and quite distinct form; why, to adventure tour travel agency perplexity of naturalists, has nature condescended to adventure tour travel agency tricks of adventure tour travel agency stage? Mr. Bates has, no doubt, hit on adventure tour travel agency true explanation. The mocked forms, which always abound in numbers, must habitually escape destruction to a large extent, otherwise they coul
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